Alcobaça Formation
Alcobaça Formation | |
---|---|
Stratigraphic range: Kimmeridgian ~156.9–152.5 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓ | |
Type | Geological formation |
Sub-units | Camadas de Guimarota |
Underlies | Lourinhã Formation |
Overlies | Montejunto Formation |
Lithology | |
Primary | Marl |
Other | Mudstone, coal, siltstone, limestone, sandstone |
Location | |
Coordinates | ![]() ![]() Alcobaça Formation (Portugal) |
The Alcobaça Formation, previously known as the Guimarota Formation and also known as the Consolação Unit,[1] is a geological formation in Portugal.[2] It dates back to the Kimmeridgian stage of the Late Jurassic. It is an important source of information on the diversity of Late Jurassic mammals.[3][4] Many of the fossils were collected from the now disused and flooded Camadas de Guimarota coal mine.
Vertebrate paleofauna
Dinosaur eggs are geographically located in Lisbon District, Portugal.[5] Dinosaur tracks are geographically located in Leiria District, Portugal.[6]
Amphibians
Amphibians of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Celtedens | Indeterminate | Porto Pinheiro | Albanerpetontid |
Choristoderes
Choristoderes of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Cteniogenys | Indeterminate | Guimarota |
Ornithischians
Indeterminate euornithopod remains located in Lisbon District.[5] Indeterminate stegosaurid remains present in Lisbon District.[5]
Ornithischians of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | District | Stratigraphic position | Abundance | Notes | Images |
Dacentrurus[5] | D. armatus[5] | Lisboa[5] | ![]() | |||
Ornithopoda[5] | Indeterminate[5] | Lisboa[5] | ||||
Stegosaurus[7] | S. cf. ungulatus | "...one tooth, five cervical vertebrae (including the axis), five dorsal vertebrae, cervical and dorsal ribs, three caudal vertebrae, chevrons, preacetabular process of the left ilium, right tibia, fibula, astragalus and [calcaneus], a cervical plate, and several fragments of plates." | ![]() | |||
Trimucrodon[5] | T. cuneatus[5] | Lisboa[5] | Later determined to be indeterminate ornithischian remains.[5] |
Saurischians
Indeterminate sauropod remains located in Leiria and Lisboa.[8]
Saurischians of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | District | Stratigraphic position | Abundance | Notes | Images |
Allosaurus[6] | Indeterminate[6] | Leiria[6] | ![]() | |||
Aviatyrannis | A. jurassica | Guimarota | ||||
Lusotitan[5] | L. atalaiensis[5] | Lisboa[5] | ![]() | |||
Ceratosaurus[5] | Indeterminate[5] | Lisboa[5] | ![]() | |||
Dinheirosaurus[5] | D. lourinhanensis[5] | Lisboa[5] | "[Two] cervicals, [nine] dorsals, ribs, limb elements."[9] | ![]() | ||
Lourinhasaurus[5] | L. alenquerensis[5] | Lisboa[5] | ||||
Megalosaurus[5] | M. insignis[5] | Lisboa[5] | Later determined to be indeterminate theropod remains.[5] |
Turtles
Turtles of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Peltochelys | P. duchastelli | Guimarota | ||||
Platychelys | Indeterminate | Guimarota | ||||
Pleurosternidae | Indeterminate | Guimarota |
Lepidosauromorphs
Lepidosauromorphs of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Becklesius | B. hoffstetteri | Guimarota | Paramacellodid lizard | |||
Dorsetisaurus | D. purbeckensis | Guimarota | Anguimorph lizard | |||
Marmoretta | Indeterminate | Guimarota | Basal Lepidosauromorph | ![]() | ||
Portugalophis | P. lignites | Guimarota | Stem-snake | |||
Saurillodon | S. proraformis | Guimarota | A lizard | |||
Saurillus | S. henkeli, cf. obtusus | Guimarota | A lizard | |||
cf. Opisthias[10] | Indeterminate | Andrés, Pombal | A rhynchocephalian |
Crocodyliformes
Crocodyliformes of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Goniopholis | G. baryglyphaeus | Guimarota | A goniopholidid | ![]() | ||
Knoetschkesuchus | K. guimarotae | Guimarota | An atoposaurid | ![]() | ||
Lisboasaurus | L. mitracostatus | Guimarota | ||||
Machimosaurus | M. hugii | Guimarota | A marine machimosaurid | ![]() |
Mammals
Mammals of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Bathmochoffatia | B. hapax | Guimarota | A paulchoffatiid multituberculate | |||
Bernardodon | B. sp, B. atlanticus | Porto Pinheiro | A pinheirodontid multituberculate | |||
Drescheratherium | D. acutum | Guimarota | A paurodontid dryolestidan | |||
Dryolestes | D. leirensis | Guimarota | A dryolestid dryolestidan | |||
Ecprepaulax | E. anomala | Porto Pinheiro | A pinheirodontid multituberculate | |||
Guimarotodon | G. leiriensis | Guimarota | A paulchoffatiid multituberculate | |||
Guimarotodus | G. inflatus | Guimarota | A dryolestid dryolestidan | |||
Haldanodon | H. exspectatus | Guimarota | A docodont | |||
Henkelodon | H. naias, H. sp | Guimarota | A paulchoffatiid multituberculate | |||
Henkelotherium | H. guimarotae | Guimarota | A dryolestidan | |||
Iberodon | I. quadrituberculatus | Porto Pinheiro | A pinheirodontid multituberculate | |||
Kielanodon | K. hopsoni | Guimarota | A paulchoffatiid multituberculate | |||
Krebsotherium | K. lusitanicum | Guimarota | A dryolestid dryolestidan | |||
Kuehneodon | K. guimarotensis, K. dietrichi, K. simpsoni, K. uniradiculatus, K. sp | Guimarota | A paulchoffatiid multituberculate | |||
Laolestes | L. andresi | Porto Pinheiro | A dryolestid dryolestidan | |||
Meketibolodon | M. robustus | Guimarota | A paulchoffatiid multituberculate | |||
Meketichoffatia | M. krausei, M. sp | Guimarota | A paulchoffatiid multituberculate | |||
Nanolestes | N. krusati | Porto Pinheiro | A cladotherian | |||
N. drescherae | Guimarota | |||||
Paulchoffatia | P. delgadoi | Guimarota | A paulchoffatiid multituberculate | |||
Pinheirodon | P. sp, P. vastus, P. pygmaeus | Porto Pinheiro | A pinheirodontid multituberculate | |||
Plesiochoffatia | P. thoas, P. peperethos | Guimarota | A paulchoffatiid multituberculate | |||
Portopinheirodon | P. asymmetricus | Porto Pinheiro | A dryolestid dryolestidan | |||
Priacodon | Indeterminate | Porto Pinheiro | A triconodontid | |||
Proalbionbaatar | P. plagiocyrtus | Guimarota | A multituberculate of uncertain placement | |||
Pseudobolodon | P. dryas, P krebsi, P. oreas | Guimarota | A paulchoffatiid multituberculate | |||
Renatodon | R. amalthea | Guimarota | A paulchoffatiid multituberculate | |||
? Tinodon | Indeterminate | Porto Pinheiro | A tinodontid symmetrodont | |||
Xenachoffatia | X. oinopion | Guimarota | A paulchoffatiid multituberculate | |||
Paulchoffatiidae | Indeterminate | Guimarota | 4 distinct genera with 7 species | |||
Pinheirodontidae | Indeterminate | Porto Pinheiro |
Color key
| Notes Uncertain or tentative taxa are in small text; |
See also
References
- ^ Mateus, O.; Dinis, J.; Cunha, P. P. (2017-09-28). "The Lourinhã Formation: the Upper Jurassic to lower most Cretaceous of the Lusitanian Basin, Portugal – landscapes where dinosaurs walked". Ciências da Terra / Earth Sciences Journal. 19 (1): 75–97. doi:10.21695/cterra/esj.v19i1.355. hdl:10316/79879. ISSN 2183-4431.
- ^ Fürsich, Franz T.; Schneider, Simon; Werner, Winfried; Lopez-Mir, Berta; Pierce, Colm S. (2021-09-13). "Life at the continental–marine interface: palaeoenvironments and biota of the Alcobaça Formation (Late Jurassic, Central Portugal), with a formal definition of the unit appended". Palaeobiodiversity and Palaeoenvironments. 102 (2): 265–329. doi:10.1007/s12549-021-00496-x. ISSN 1867-1608.
- ^ "Guimarota". Paleobiology Database. Retrieved 31 August 2018.
- ^ "Porto Pinheiro, Lourinha". Paleobiology Database. Retrieved 31 August 2018.
- ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
- ^ a b c d "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
- ^ Escaso, Fernando; Ortega, Francisco; Dantas, Pedro; Malafaia, Elisabete; Pimentel, Nuno L.; Pereda-Suberbiola, Xabier; Sanz, José Luis; Kullberg, José Carlos; Kullberg, María Carla (2006-12-23). "New evidence of shared dinosaur across Upper Jurassic Proto-North Atlantic: Stegosaurus from Portugal". Naturwissenschaften. 94 (5): 367–374. Bibcode:2007NW.....94..367E. doi:10.1007/s00114-006-0209-8. PMID 17187254. S2CID 10930309.
- ^ "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," and "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Pages 548-549.
- ^ "Table 13.1," in Weishampel, et al. (2004). Page 265.
- ^ Malafaia, E.; Ortega, F.; Escaso, F.; Dantas, P.; Pimentel, N.; Gasulla, J. M.; Ribeiro, B.; Barriga, F.; Sanz, J. L. (2010). "Vertebrate fauna at the Allosaurus fossil-site of Andrés (Upper Jurassic), Pombal, Portugal". Journal of Iberian Geology. 36 (2): 193–204. doi:10.5209/rev_JIGE.2010.v36.n2.7. ISSN 1698-6180.
Bibliography
- Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (2004), The Dinosauria, 2nd edition, Berkeley: University of California Press, pp. 1–880, ISBN 0-520-24209-2, retrieved 2019-02-21