Family of beetles
Helotidae
Temporal range: Aptian–Recent PreꞒ Ꞓ O S D C P T J K Pg N |
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Neohelota sp. |
Scientific classification |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Coleoptera |
Suborder: | Polyphaga |
Infraorder: | Cucujiformia |
Superfamily: | Cucujoidea |
Family: | Helotidae Chapuis, 1876 |
Genera |
- Helota
- Neohelota
- Afrohelotina
- Metahelotella
- Strophohelota
- †Palaeohelota
- †Metahelotella
- †Trihelota
- †Burmahelota
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Helotidae is a family of beetles, in the suborder Polyphaga. The family includes about five extant genera, Helota MacLeay, Neohelota Ohta, Afrohelotina Kirejtshuk, Metahelotella Kirejtshuk, and Strophohelota Kirejtshuk.[1][2] Helotidae are found mainly in the Old World tropics and are absent from Australia and Madagascar. The antennae are clubbed on the final three segments and is retractable within grooves under the head. The wings have reduced venation with just 4 anal veins.[3] Helotids are known to be associated with sap, fruit and flowers, and the larvae of some species are known to bore into wood in order to pupate.[4]
A few fossil genera have been described including Palaeohelota from the Lower Cretaceous (Aptian) Yixian Formation in China, Burmahelota, Metahelotella and Trihelota from the Upper Cretaceous (Cenomanian) Burmese amber of Northern Myanmar. Another fossil genus Laodiscis placed in this family originally is now considered of uncertain taxonomic position.[5]
References
- ^ Lee, Chi-Feng; Votruba, Petr (2013). "A revision of the family Helotidae (Coleoptera: Cucujoidea): VIII. The guerinii species-group of the genus Neohelota". Entomologica Basiliensia et Collectionis Frey. 34: 269–308.
- ^ Lee, Chi-Feng (2007). "Revision of Family Helotidae (Coleoptera: Cucujoidea):I. Gemmata Group of Genus Helota". Ann. Entomol. Soc. Am. 100 (5): 623–639.
- ^ Crowson, R.A. (1967). The natural classification of the families of Coleoptera. Middlesex: E.W. Classey Ltd. p. 104.
- ^ Lawrence, John F.; Ślipinśki, Adam; Lee, Chi-Feng (2010-12-31), Kükenthal, Willy; Leschen, Richard A.B.; Beutel, Rolf G.; Lawrence, John F. (eds.), "10.3. Helotidae Reitter, 1876/Chapuis, 1876", Coleoptera, Beetles, Volume 2, Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim), DE GRUYTER, pp. 292–295, doi:10.1515/9783110911213.292, ISBN 978-3-11-019075-5, retrieved 2022-10-02
- ^ Liu, Zhenhua; Ślipiński, Adam; Ren, Dong; Pang, Hong (2019). "The first Mesozoic Helotidae (Coleoptera: Cucujoidea)". Cretaceous Research. 96: 113–119. doi:10.1016/j.cretres.2018.12.005.
Additional sources
- Ritsema C. (1915) A systematic catalogue of the Coleopterous family Helotidae in the Leiden Museum Zoologische Mededelingen, Vol. 1 P. 125-139 PDF
- Wegrzynowicz P. (2000) Catalogue of the Helotidae (Coleoptera: Cucujoidea), Zoologische Mededelingen, Vol. 73 P. 391-411 PDF
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- Crowsoniellidae (Crowsoniella relicta)
- Cupedidae (reticulated beetles)
- Jurodidae (Sikhotealinia zhiltzovae)
- Micromalthidae
- Ommatidae
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Extant families | - Amphizoidae (trout-stream beetles)
- Aspidytidae
- Carabidae (ground beetles)
- Cicindelidae (tiger beetles)
- Dytiscidae (predaceous diving beetles)
- Gyrinidae (whirligig beetles)
- Haliplidae (crawling water beetles)
- Hygrobiidae
- Meruidae (Meru phyllisae)
- Noteridae (burrowing water beetles)
- Trachypachidae (false ground beetles)
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Bostrichiformia | Bostrichoidea | - Bostrichidae (auger beetles)
- Dermestidae (skin beetles)
- Endecatomidae
- Jacobsoniidae (Jacobson's beetles)
- Nosodendridae (wounded-tree beetles)
- Ptiniidae (furniture beetles, death watch beetles, spider beetles)
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Derodontoidea | - Derodontidae (tooth-necked fungus beetles)
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Cucujiformia | Chrysomeloidea | |
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Cleroidea | |
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Coccinelloidea | |
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Cucujoidea | |
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Curculionoidea (weevils) | - Anthribidae (fungus weevils)
- Attelabidae (leaf-rolling weevils)
- Belidae (primitive weevils)
- Brentidae (straight snout weevils, New York weevil)
- Caridae
- Curculionidae (true weevils, bark beetles, ambrosia beetles)
- Nemonychidae (pine flower weevils)
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Lymexyloidea | - Lymexylidae (ship-timber beetles)
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Tenebrionoidea | - Aderidae (ant-like leaf beetles)
- Anthicidae (ant-like flower beetles)
- Archeocrypticidae (cryptic fungus beetles)
- Boridae (conifer bark beetles)
- Chalcodryidae
- Ciidae (minute tree-fungus beetles)
- Melandryidae (false darkling beetles)
- Meloidae (blister beetles)
- Mordellidae (tumbling flower beetles)
- Mycetophagidae (hairy fungus beetles)
- Mycteridae (palm and flower beetles)
- Oedemeridae (false blister beetle)
- Perimylopidae, or Promecheilidae
- Prostomidae (jugular-horned beetles)
- Pterogeniidae
- Pyrochroidae (fire-coloured beetles)
- Pythidae (dead log bark beetles)
- Ripiphoridae (wedge-shaped beetles)
- Salpingidae (narrow-waisted bark beetles)
- Scraptiidae (false flower beetles)
- Stenotrachelidae (false longhorn beetles)
- Synchroidae (synchroa bark beetles)
- Tenebrionidae (darkling beetles)
- Tetratomidae (polypore fungus beetles)
- Trictenotomidae
- Ulodidae
- Zopheridae (ironclad beetles, cylindrical bark beetles)
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Elateriformia | Buprestoidea | - Buprestidae (jewel beetles, or metallic wood-boring beetles)
- Schizopodidae
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Byrrhoidea | - Byrrhidae (pill beetles)
- Callirhipidae (cedar beetles)
- Chelonariidae (turtle beetles)
- Cneoglossidae
- Dryopidae (long-toed water beetles)
- Elmidae (riffle beetles)
- Eulichadidae (forest stream beetles)
- Heteroceridae (variegated mud-loving beetles)
- Limnichidae (minute mud beetles)
- Lutrochidae (travertine beetles)
- Psephenidae (water-penny beetles)
- Ptilodactylidae
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Dascilloidea | - Dascillidae (soft bodied plant beetles)
- Rhipiceridae (cicada beetle, cicada parasite beetles)
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Elateroidea | - Artematopodidae (soft-bodied plant beetles)
- Brachypsectridae (Texas beetles)
- Cantharidae (soldier beetles)
- Cerophytidae (rare click beetles)
- Elateridae (click beetles)
- Eucnemidae (false click beetles)
- Jurasaidae
- Lampyridae (fireflies)
- Lycidae (net-winged beetles)
- Omethidae (false fireflies, long-lipped beetles)
- Phengodidae (glowworm beetles)
- Rhagophthalmidae
- Sinopyrophoridae
- Throscidae (false metallic wood-boring beetles)
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Rhinorhipoidea | - Rhinorhipidae (Rhinorhipus tamborinensis)
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Scirtoidea | |
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Scarabaeiformia | Scarabaeoidea | - Belohinidae (Belohina inexpectata)
- Bolboceratidae
- Diphyllostomatidae (false stag beetles)
- Geotrupidae (dor beetles)
- Glaphyridae (bumble bee scarab beetles)
- Glaresidae (enigmatic scarab beetles)
- Hybosoridae (scavenger scarab beetles)
- Lucanidae (stag beetles)
- Ochodaeidae (sand-loving scarab beetles)
- Passalidae (betsy beetles)
- Pleocomidae (rain beetles)
- Scarabaeidae (scarabs)
- Trogidae (hide beetles)
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Staphyliniformia | |
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Taxon identifiers |
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Helotidae | |
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