South American fox

Genus of carnivores

Lycalopex[1]
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Family: Canidae
Tribe: Canini
Subtribe: Cerdocyonina
Genus: Lycalopex
Burmeister 1854
Type species
Canis magellanicus[3]
Species
  • Lycalopex culpaeus
  • Lycalopex fulvipes
  • Lycalopex griseus
  • Lycalopex gymnocercus
  • Lycalopex sechurae
  • Lycalopex vetulus
  • Canis (Pseudalopex) australis (Kerr 1792)[2]
Range of the six living zorro species
Synonyms
  • Pseudalopex Burmeister 1856
  • Canis (Pseudalopex) Allen 1895

The South American foxes (Lycalopex), commonly called raposa in Portuguese, or zorro in Spanish, are a genus from South America of the subfamily Caninae. Despite their name, they are not true foxes, but are a unique canid genus more closely related to wolves and jackals than to true foxes; some of them resemble foxes due to convergent evolution. The South American gray fox, Lycalopex griseus, is the most common species, and is known for its large ears and a highly marketable, russet-fringed pelt.

The second-oldest known fossils belonging to the genus were discovered in Chile, and date from 2.0 to 2.5 million years ago, in the mid- to late Pliocene.[4] The Vorohué Formation of Argentina has provided older fossils, dating to the Uquian to Ensenadan (Late Pliocene).[5]

Names

The common English word "zorro" is a loan word from Spanish, with the word originally meaning "fox". Current usage lists Pseudalopex (literally: "false fox") as synonymous with Lycalopex ("wolf fox"), with the latter taking precedence.[1][6] In 1895, Allen classified Pseudalopex as a subgenus of Canis, establishing the combination Canis (Pseudalopex), a name still used in the fossil record.[2]

Species

Species currently included in this genus include:[1]

Image Name Common name Distribution
Lycalopex culpaeus Culpeo or Andean fox Ecuador and Peru to the southern regions of Patagonia and Tierra del Fuego
Lycalopex fulvipes Darwin's fox Nahuelbuta National Park (Araucanía Region), the Valdivian Coastal Range (Los Ríos Region) in mainland Chile and Chiloé Island
Lycalopex griseus South American gray fox or chilla Argentina and Chile
Lycalopex gymnocercus Pampas fox northern and central Argentina, Uruguay, eastern Bolivia, Paraguay, and southern Brazil
Lycalopex sechurae Sechuran fox west-central, northwestern Peru, including the Sechura Desert, and southwestern Ecuador
Lycalopex vetulus Hoary fox south-central Brazil
Canis (Pseudalopex) australis Vorohué Formation, Uquian-Ensenadan Argentina[5]

In 1914, Oldfield Thomas established the genus Dusicyon, in which he included these zorros. They were later reclassified to Lycalopex (via Pseudalopex) by Langguth in 1975.[1]

Phylogeny

The following phylogenetic tree shows the evolutionary relationships between the Lycalopex species, based on molecular analysis of mitochondrial DNA control region sequences.[7]

Relationship with humans

The zorros are hunted in Argentina for their durable, soft pelts. They are also often labelled 'lamb-killers'.[citation needed]

In his diary of his well-known 1952 traveling with the young Che Guevara,[8] Alberto Granado mentions talking with seasonal workers employed on vast sheep farms, who told him of a successful campaign by the ranch owners to exterminate the foxes who were preying on lambs. The ranchers offered a reward of one Argentinian peso for the body of a dead male fox and as much as five pesos for a female fox; to impoverished workers in the early 1950s, five pesos were a significant sum. Within a few years, foxes became virtually extinct in a large part of Argentina.

The Fuegian dog (Spanish: perro yagán, perro fueguino), also known as the Yaghan dog, was a domesticated form of the culpeo (Lycalopex culpaeus),[9] unlike other domesticated canids which were dogs and silver foxes. This means different canid species have been domesticated multiple times by humans independently.

References

  1. ^ a b c d Wozencraft, W. C. (2005). "Order Carnivora". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. ISBN 978-0-8018-8221-0. OCLC 62265494.
  2. ^ a b "Canis (Pseudalopex) australis Kerr 1792". Fossilworks. Gateway to the Paleobiology Database. Retrieved 15 December 2021.
  3. ^ Wilson, D. E.; Reeder, D. M., eds. (2005). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. ISBN 978-0-8018-8221-0. OCLC 62265494.
  4. ^ Lucherini, M. & Luengos Vidal, E. M. (2008). "Lycalopex gymnocercus (Carnivora: Canidae)". Mammalian Species. 820: Number 820, pp. 1–9. doi:10.1644/820.1. hdl:11336/115900.
  5. ^ a b Vorohuen (sic; Vorohué) Formation at Fossilworks.org
  6. ^ Lucherini, M. (2016). "Lycalopex culpaeus". IUCN Red List of Threatened Species. 2016: e.T6929A85324366. doi:10.2305/IUCN.UK.2016-1.RLTS.T6929A85324366.en. Retrieved 11 November 2021.
  7. ^ Tchaicka, Ligia; Freitas, Thales Renato Ochotorena de; Bager, Alex; Vidal, Stela Luengos; Lucherini, Mauro; Iriarte, Agustín; Novaro, Andres; Geffen, Eli; Garcez, Fabricio Silva; Johnson, Warren E.; Wayne, Robert K.; Eizirik, Eduardo (2016). "Molecular assessment of the phylogeny and biogeography of a recently diversified endemic group of South American canids (Mammalia: Carnivora: Canidae)" (PDF). Genetics and Molecular Biology. 39 (3): 442–451. doi:10.1590/1678-4685-GMB-2015-0189. PMC 5004827. PMID 27560989.
  8. ^ TRAVELLING WITH CHE GUEVARA by Alberto Granado
  9. ^ Petrigh, Romina S.; Fugassa, Martin H. (December 13, 2013). "Molecular identification of a Fuegian dog belonging to the Fagnano Regional Museum ethnographic collection, Tierra del Fuego" (PDF). Quaternary International. 317: 14–18. Bibcode:2013QuInt.317...14P. doi:10.1016/j.quaint.2013.07.030. Archived from the original (PDF) on December 20, 2016. Retrieved September 2, 2020.

Further reading

Wikimedia Commons has media related to Pseudalopex.
Wikispecies has information related to Lycalopex.
  • Nowak, Ronald M. (2005). Walker's Carnivores of the World. Baltimore: Johns Hopkins Press. ISBN 0-8018-8032-7
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Extant Carnivora species
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Viverroidea
    • see below↓
Hemigalinae
  • Owston's palm civet (Chrotogale owstoni)
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Paradoxurinae
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Paradoxurus
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Viverrinae
sensu lato
Viverrinae
sensu stricto
  • Small Indian civet (Viverricula indica)
  • African civet (Civettictis civetta)
Viverra
  • Malabar large-spotted civet (V. civettina)
  • Large-spotted civet (V. megaspila)
  • Malayan civet (V. tangalunga)
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Genettinae
Poiana
(African linsangs)
  • Central African oyan (P. richardsonii)
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Genetta
(genets)
  • Abyssinian genet (G. abyssinica)
  • Angolan genet (G. angolensis)
  • Bourlon's genet (G. bourloni)
  • Crested servaline genet (G. cristata)
  • Common genet (G. genetta)
  • Johnston's genet (G. johnstoni)
  • Letaba genet (G. letabae)
  • Rusty-spotted genet (G. maculata)
  • Pardine genet (G. pardina)
  • Aquatic genet (G. piscivora)
  • King genet (G. poensis)
  • Servaline genet (G. servalina)
  • Hausa genet (G. thierryi)
  • Cape genet (G. tigrina)
  • Giant forest genet (G. victoriae)
  • South African small-spotted genet (G. felina)
Herpestoidea
    • see below↓
Hyaenidae
(hyenas)
Proteles
  • Aardwolf (P. cristata)
Hyaeninae
(bone-crushing hyenas)
  • Striped hyena (Hyaena hyaena)
  • Brown hyena (Parahyaena brunnea)
Crocuta
  • Spotted hyena (C. crocuta)
Herpestidae sensu lato
Eupleridae
(Malagasy
carnivorans)
Euplerinae
  • Fossa (Cryptoprocta ferox)
  • Malagasy civet (Fossa fossana)
Eupleres
  • Eastern falanouc (E. goudotii)
  • Western falanouc (E. major)
Galidiinae
  • Ring-tailed vontsira (Galidia elegans)
Galidictis
  • Broad-striped Malagasy mongoose (G. fasciata)
  • Grandidier's mongoose (G. grandidieri)
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Salanoia
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  • Durrell's vontsira (S. durrelli)
Herpestidae
sensu stricto

(mongooses)
Mungotinae
Suricata
  • Meerkat (S. suricatta)
  • Liberian mongoose (Liberiictus kuhni)
Mungos
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  • Pousargues's mongoose (Dologale dybowskii)
Helogale
  • Ethiopian dwarf mongoose (H. hirtula)
  • Common dwarf mongoose (H. parvula)
Crossarchus
  • Alexander's kusimanse (C. alexandri)
  • Angolan kusimanse (C. ansorgei)
  • Common kusimanse (C. obscurus)
  • Flat-headed kusimanse (C. platycephalus)
Herpestinae
  • Marsh mongoose (Atilax paludinosus)
  • Long-nosed mongoose (Xenogale naso)
Urva
  • Small Indian mongoose (U. auropunctata)
  • Short-tailed mongoose (U. brachyura)
  • Indian grey mongoose (U. edwardsii)
  • Indian brown mongoose (U. fusca)
  • Javan mongoose (U. javanica)
  • Collared mongoose (U. semitorquata)
  • Ruddy mongoose (U. smithii)
  • Crab-eating mongoose (U. urva)
  • Stripe-necked mongoose (U. vitticolla)
  • White-tailed mongoose (Ichneumia albicauda)
  • Yellow mongoose (Cynictis penicillata)
  • Selous's mongoose (Paracynictis selousi)
  • Meller's mongoose (Rhynchogale melleri)
Bdeogale
  • Bushy-tailed mongoose (B. crassicauda)
  • Jackson's mongoose (B. jacksoni)
  • Black-footed mongoose (B. nigripes)
Herpestes
  • Angolan slender mongoose (H. flavescens)
  • Egyptian mongoose (H. ichneumon)
  • Somalian slender mongoose (H. ochracea)
  • Cape gray mongoose (H. pulverulenta)
  • Common slender mongoose (H. sanguinea)
Caniformia ("dog-like" carnivorans)
Canidae (dogs)
Urocyon
  • Gray fox (U. cinereoargenteus)
  • Island fox (U. littoralis)
Vulpini
  • Bat-eared fox (Otocyon megalotis)
Nyctereutes
  • Common raccoon dog (N. procyonoides)
  • Japanese raccoon dog (N. viverrinus)
Vulpes
(foxes)
  • Bengal fox (V. bengalensis)
  • Blanford's fox (V. cana)
  • Cape fox (V. chama)
  • Corsac fox (V. corsac)
  • Tibetan fox (V. ferrilata)
  • Arctic fox (V. lagopus)
  • Kit fox (V. macrotis)
  • Pale fox (V. pallida)
  • Rüppell's fox (V. rueppelli)
  • Swift fox (V. velox)
  • Red fox (V. vulpes)
  • Fennec fox (V. zerda)
Canini
(true dogs)
Cerdocyonina
(zorro)
  • Maned wolf (Chrysocyon brachyurus)
Speothos
  • Bush dog (S. venaticus)
  • Short-eared dog (Atelocynus microtis)
  • Crab-eating fox (Cerdocyon thous)
Lycalopex
  • Culpeo (L. culpaeus)
  • Darwin's fox (L. fulvipes)
  • South American gray fox (L. griseus)
  • Pampas fox (L. gymnocercus)
  • Sechuran fox (L. sechurae)
  • Hoary fox (L. vetulus)
Canina
(wolf-like canids)
Lupulella
  • Side-striped jackal (L. adustus)
  • Black-backed jackal (L. mesomelas)
  • African wild dog (Lycaon pictus)
  • Dhole (Cuon alpinus)
Canis
  • Golden jackal (C. aureus)
  • Domestic dog (C. familiaris)
  • Coyote (C. latrans)
  • African wolf (C. lupaster)
  • Wolf (C. lupus)
  • Eastern wolf (C. lycaon)
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  • Ethiopian wolf (C. simensis)
Ursidae
(bears)
Ailuropoda
  • Giant panda (A. melanoleuca)
Tremarctos
  • Spectacled bear (T. ornatus)
Ursinae
  • Sloth bear (Melursus ursinus)
  • Sun bear (Helarctos malayanus)
Ursus
  • American black bear (U. americanus)
  • Asian black bear (U. thibetanus)
  • Brown bear (U. arctos)
  • Polar bear (U. maritimus)
Mustelida
Pinnipedia (seals)
    • see below↓
Musteloidea
    • see below↓
Pinnipedia (seals)
Odobenidae
  • Walrus (Odobenus rosmarus)

Otariidae
(eared seals:
fur seals,
sea lions)
Callorhinus
  • Northern fur seal (C. ursinus)
Otariinae
  • Steller sea lion (Eumetopias jubatus)
Zalophus
  • California sea lion (Z. californianus)
  • Galápagos sea lion (Z. wollebaeki)
  • South American sea lion (Otaria flavescens)
Neophoca
  • Australian sea lion (N. cinerea)
  • New Zealand sea lion (Phocarctos hookeri)
Arctocephalus
  • South American fur seal (A. australis)
  • Australasian fur seal (A. forsteri)
  • Galápagos fur seal (A. galapagoensis)
  • Antarctic fur seal (A. gazella)
  • Juan Fernández fur seal (A. philippii)
  • Brown fur seal (A. pusillus)
  • Guadalupe fur seal (A. townsendi)
  • Subantarctic fur seal (A. tropicalis)
Phocidae
(earless seals
or true seals)
Phocinae
("northern seals")
  • Bearded seal (Erignathus barbatus)
  • Hooded seal (Cystophora cristata)
Phocini
  • Harp seal (Pagophilus groenlandicus)
  • Ribbon seal (Histriophoca fasciata)
  • Grey seal (Halichoerus grypus)
Phoca
  • Spotted seal (P. largha)
  • Harbor seal (P. vitulina)
Pusa
  • Caspian seal (P. caspica)
  • Ringed seal (P. hispida)
  • Baikal seal (P. sibirica)
Monachinae
("southern seals")
Monachini
(monk seals)
  • Mediterranean monk seal (Monachus monachus)
Neomonachus
  • Hawaiian monk seal (N. schauinslandi)
Mirounga
(elephant seals)
  • Northern elephant seal (M. angustirostris)
  • Southern elephant seal (M. leonina)
Lobodontini
(Antarctic seals)
  • Leopard seal (Hydrurga leptonyx)
  • Weddell seal (Leptonychotes weddellii)
  • Crabeater seal (Lobodon carcinophagus)
  • Ross seal (Ommatophoca rossi)
Ailuridae
  • Red panda (Ailurus fulgens)
Mephitidae
(skunks)
Conepatus
(hog-nosed skunks)
  • Molina's hog-nosed skunk (C. chinga)
  • Humboldt's hog-nosed skunk (C. humboldtii)
  • American hog-nosed skunk (C. leuconotus)
  • Striped hog-nosed skunk (C. semistriatus)
Mephitis
  • Hooded skunk (M. macroura)
  • Striped skunk (M. mephitis)
Mydaus
(stink badgers)
  • Sunda stink badger (M. javanensis)
  • Palawan stink badger (M. marchei)
Spilogale
(spotted skunks)
  • Southern spotted skunk (S. angustifrons)
  • Western spotted skunk (S. gracilis)
  • Eastern spotted skunk (S. putorius)
  • Pygmy spotted skunk (S. pygmaea)
Procyonidae
  • Kinkajou (Potos flavus)
Bassariscus
  • Ringtail (B. astutus)
  • Cacomistle (B. sumichrasti)
Procyon
(raccoons)
  • Crab-eating raccoon (P. cancrivorus)
  • Raccoon (P. lotor)
  • Cozumel raccoon (P. pygmaeus)
Bassaricyon
(olingos)
  • Eastern lowland olingo (B. alleni)
  • Northern olingo (B. gabbii)
  • Western lowland olingo (B. medius)
  • Olinguito (B. neblina)
Nasuina
(coatis)
Nasua
  • South American coati (N. nasua)
  • White-nosed coati (N. narica)
Nasuella
  • Eastern mountain coati (N. meridensis)
  • Western mountain coati (N. olivacea)
Mustelidae
    • see below↓
  • American badger (Taxidea taxus)
Mellivora
  • Honey badger (M. capensis)
Melinae
(Eurasian badgers)
Arctonyx
  • Northern hog badger (A. albogularis)
  • Greater hog badger (A. collaris)
  • Sumatran hog badger (A. hoevenii)
Meles
  • Japanese badger (M. anakuma)
  • Caucasian badger (M. canescens)
  • Asian badger (M. leucurus)
  • European badger (M. meles)
Melogale
(ferret-badgers)
  • Vietnam ferret-badger (M. cucphuongensis)
  • Bornean ferret badger (M. everetti)
  • Chinese ferret-badger (M. moschata)
  • Javan ferret-badger (M. orientalis)
  • Burmese ferret-badger (M. personata)
  • Formosan ferret-badger (M. subaurantiaca)
Guloninae
  • Tayra (Eira barbara)
Pekania
  • Fisher (P. pennanti)
Gulo
  • Wolverine (G. gulo)
Martes
(martens)
  • American marten (M. americana)
  • Pacific marten (M. caurina)
  • Yellow-throated marten (M. flavigula)
  • Beech marten (M. foina)
  • Nilgiri marten (M. gwatkinsii)
  • European pine marten (M. martes)
  • Japanese marten (M. melampus)
  • Sable (M. zibellina)
Ictonychinae
Lyncodontini
  • Patagonian weasel (Lyncodon patagonicus)
Galictis
(grisons)
  • Lesser grison (G. cuja)
  • Greater grison (G. vittata)
Ictonychini
(African polecats)
Vormela
  • Marbled polecat (V. peregusna)
  • African striped weasel (Poecilogale albinucha)
Ictonyx
  • Saharan striped polecat (I. libycus)
  • Striped polecat (I. striatus)
Lutrinae
(otters)
  • Giant otter (Pteronura brasiliensis)
Lontra
  • North American river otter (L. canadensis)
  • Marine otter (L. felina)
  • Neotropical otter (L. longicaudis)
  • Southern river otter (L. provocax)
Enhydra
  • Sea otter (E. lutris)
  • Spotted-necked otter (Hydrictis maculicollis)
Lutra
  • Eurasian otter (L. lutra)
  • Hairy-nosed otter (L. sumatrana)
Lutrogale
  • Smooth-coated otter (L. perspicillata)
Aonyx
  • African clawless otter (A. capensis)
  • Asian small-clawed otter (A. cinereus)
  • Congo clawless otter (A. congicus)
Mustelinae
Neogale
  • Amazon weasel (N. africana)
  • Colombian weasel (N. felipei)
  • Long-tailed weasel (N. frenata)
  • American mink (N. vison)
Mustela
(weasels)
  • Sichuan weasel (M. aistoodonnivalis)
  • Mountain weasel (M. altaica)
  • Stoat/Beringian ermine (M. erminea)
  • Steppe polecat (M. eversmannii)
  • Ferret (M. furo)
  • Haida ermine (M. haidarum)
  • Japanese weasel (M. itatsi)
  • Yellow-bellied weasel (M. kathiah)
  • European mink (M. lutreola)
  • Indonesian mountain weasel (M. lutreolina)
  • Black-footed ferret (M. nigripes)
  • Least weasel (M. nivalis)
  • Malayan weasel (M. nudipes)
  • European polecat (M. putorius)
  • American ermine (M. richardsonii)
  • Siberian weasel (M. sibirica)
  • Back-striped weasel (M. strigidorsa)
Taxon identifiers
Lycalopex
Pseudalopex
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